Department of Ophthalmology, Graduate School of Medical Sciences, Kyushu University, Fukuoka, Japan. Department of Ophthalmology, Graduate School of Medical Sciences, Kyushu University, Fukuoka, Japan. snakao@med.kyushu-u.ac.jp. Department of Ophthalmology, National Kyushu Medical Center, 1-8-1, Jigyo-hama, Chuo-ku, Fukuoka, 8108563, Japan. snakao@med.kyushu-u.ac.jp. Department of Ophthalmology, Fukuoka University Chikushi Hospital, Fukuoka, Japan. Centre for Experimental Medicine, Queen's University Belfast, Belfast, Northern Ireland.